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My CV can be viewed here.  Reprints can be found through the links below, Google Scholar, ResearchGate, or upon request.

Edited Books

Gilbert CC and Frost SR, eds.  (under contract).  The Fossil Cercopithecidae of the African Plio-Pleistocene: Systematics, Evolution, and Biochronology. Springer.

Fleagle JG, Gilbert CC, and Baden AL.  (expected 2024).  Primate Adaptation and Evolution, 4th edition.  San Diego: Academic Press/Elsevier. 

Fleagle JG and Gilbert CC, eds.  (2008).  Elwyn Simons: A Search for Origins.  New York: Springer.

Peer-Reviewed Journal Articles

Sehgal RK, Singh NP, Singh AP, Gilbert CC, Patel BA, and Patnaik R.  (accepted, in press).  First report of rodents from the Miocene Siwalik locality of Dunera, Pathankot District, Punjab, India.  Palaeontologia Electronica.

*Post NW, Gilbert CC, Pugh KD, Mongle CS.  (2023).  Implications of outgroup selection in the phylogenetic inference of hominoids and fossil hominins.  Journal of Human Evolution 184: 103437.

Frost SR, White F, *Reda H, Gilbert CC.  (2022).  Biochronology of South African hominin-bearing sites: a reassessment using cercopithecid primates.  Proceedings of the National Academy of Sciences, USA 119, e2210627119.

Rowan J, Lazagabaster IA, Campisano CJ, Bibi F, Bobe R, Boisserie J-R, Frost SR, Getachew T, Gilbert CC, Lewis ME, Melaku S, Scott E, Souron A, Werdelin L, Kimbel WH, and Reed KE (in revision).  Early Pleistocene large mammals from Maka’amitalu, Hadar, lower Awash Valley, Ethiopia.  PeerJ 10, e13210.


Sehgal RK, Singh AP, Gilbert CC, Patel BA, Campisano CJ, Selig KR, Patnaik R, and Singh NP.  (2022).  First occurrence of a treeshrew and description of other micromammals from the middle Miocene hominoid locality of Ramnagar, Udhampur district, Jammu and Kashmir.  Journal of Paleontology 96: 1318-1335.  DOI:


Arenson JL, Harrison T, Sargis EJ, Taboada HG, and Gilbert CC.  (2022).  A new species of fossil guenon (Cercopithecini, Cercopithecidae) from the Early Pleistocene Lower Ngaloba Beds, Laetoli, Tanzania.  Journal of Human Evolution 163: 103136.


Delson E, Alba DM, Frost SR, Getahun DA, Gilbert CC.  (2021).  Case 3847- Simopithecus oswaldi Andrews, 1916 (currently Theropithecus oswaldi; Mammalia, Primates, Cercopithecidae), proposed conservation by reversal of precedence with Cynocephalus atlanticus Thomas, 1884.  The Bulletin of Zoological Nomenclature 78, 99-111.


Gilbert CC, Gilissen E, Arenson JL, Patel BA, Nakatsukasa M, Hart TB, Hart JA, Detwiler KM, and Sargis EJ.  (2021).  Morphological analysis of new Cercopithecus dryas specimens from the central Congo Basin: taxonomic considerations and an emended diagnosis.  American Journal of Physical Anthropology 176: 361-389.


Smaers JB, Rothman RS, Hudson DR, Balanoff AM, Beatty B, Dechmann DKN, DeVries D, Dunn JC, Fleagle JG, Gilbert CC, Goswami A, Iwaniuk AN, Jungers WL, Kerney M, Ksepka DT, Manger PR, Mongle CS, Rohlf FJ, Smith NA, Soligo C, Weisbecker V, Safi K.  (2021).  The evolution of mammalian brain size.  Science Advances 7: eabe210.


Gilbert CC, Ortiz A., Patel BA, Singh NP, Campisano CJ, Fleagle JG, *Pugh KD, and Patnaik R.  (2020).  New Middle Miocene ape (Primates: Hylobatidae) from Ramnagar, India fills major gaps in the hominoid fossil record.  Proceedings of the Royal Society B 287: 20201655.


Arenson JL, Sargis EJ, Hart JA, Hart TB, Detwiler KM, and Gilbert CC.  (2020).  Skeletal morphology of the lesula (Cercopithecus lomamiensis) and the evolution of guenon locomotor behavior.  American Journal of Physical Anthropology 172: 3-24.


Gilbert CC, Sehgal, RK, Pugh KD, Campisano CJ, Singh, NP, May E, Patel BA, Patnaik R.  (2019).  New Sivapithecus specimen from Ramnagar (J & K), India and a taxonomic revision of Ramnagar hominoids.  Journal of Human Evolution 135: 102665. (


Pugh KD and Gilbert CC.  (2018).  Phylogenetic relationships of living and fossil African papionins: combined evidence from morphology and molecules.  Journal of Human Evolution 123: 35-51.


Gilbert CC, Frost SR, Pugh KD, Anderson M, and Delson E (2018).  Evolution of the modern baboon (Papio hamadryas): A reassessment of the African Plio-Pleistocene record.  Journal of Human Evolution 122: 38-69.

Fuchs AJ, Gilbert CC, Kamilar JM.  (2018).  Ecological niche modeling of the genus PapioAmerican Journal of Physical Anthropology.  

Singh NP, Gilbert CC, Patel BA, Patnaik R.  (2018).  The taphonomy, palaeoecology and biochronology of the Middle Miocene hominoid locality of Ramnagar (Jammu and Kashmir, India).  Journal of Asian Earth Sciences.


Nengo I, Tafforeau P, Gilbert CC, Fleagle JG, Miller E, Feibel C, Fox D, Feinberg J, Pugh KD, Berruyer C, Engle Z, Spoor F.  (2017).  New infant cranium from the African Miocene sheds light on ape evolution.  Nature 548: 169-174.  


Gilbert CC, Jungers WL.  (2017).  Comment on relative brain size in early primates and the use of encephalization quotients in primate evolution.  Journal of Human Evolution 109: 79-87.


Gilbert CC, Patel BA, Singh NP, Campisano CJ, Fleagle JG, Rust KL, and Patnaik R.  (2017).  New sivaladapid primate from Lower Siwalik deposits surrounding Ramnagar, Jammu and Kashmir State, India.  Journal of Human Evolution 102: 21-41.


Singleton M, Gilbert CC, Frost SR, Seitelman, BC.  (2016).  Comparative morphometric analysis of a juvenile papionin (Primates: Cercopithecidae) from Kromdraai A.  Annals of the Ditsong Museum of Natural History 6: 1-17.  


Boyer DM, Kirk EC, Silcox MT, Gunnell GF, Gilbert CC, Yapuncich GS, Allen KL, Welch E, Bloch JI, Gonzalez L, Kay RF, Seiffert ER.  (2016).  Internal carotid arterial canal size and scaling in Euarchonta: re-assessing implications for arterial patency and phylogenetic relationships in early fossil primates.  Journal of Human Evolution 97: 123-144.


Fleagle JG, Gilbert CC, and Baden AL.  (2016).  Comparing primate crania: the importance of fossils. American Journal of Physical Anthropology 161: 259-275.


Gilbert CC, Frost SR, and Delson E.  (2016a).  Reassessment of the Olduvai Bed I cercopithecoids: a new biochronological and biogeographical link to the South African fossil record.  Journal of Human Evolution 92: 50-59.


Gilbert CC, Takahashi MQ, Delson E.  (2016b).  Cercopithecoid humeri from Taung support the distinction of major papionin clades in the South African fossil record.  Journal of Human Evolution 90: 88-104.


Gilbert CC, Steininger CM, Kibii JM, Berger LR.  (2015).  Papio cranium from the hominin-bearing site of Malapa: implications for the evolution of modern baboon cranial morphology and South African Plio-Pleistocene biochronology.  PLoS ONE 10(8), e0133361. doi:10.1371/journal.pone.0133361


DeVreese L, Gilbert CC.  (2015).  Phylogenetic relationships within the Cercocebus-Mandrillus clade as indicated by craniodental morphology: implications for evolutionary biogeography.  American Journal of Physical Anthropology 158: 227-241.


Frost SR, Gilbert CC, Pugh KD, Guthrie E, Delson E.  (2015).  The hand of Cercopithecoides williamsi (Mammalia, Primates): Earliest fossil evidence for pollical reduction among colobine monkeys and its implications for their evolution.  PLoS ONE 10(5): e0125030. doi:10.1371/journal.pone.0125030


Gilbert CC, Maiolino S.  (2015).  Comment to “Primates in the Eocene” by Gingerich (2012).  Palaeobiodiversity and Palaeoenvironments 95: 237-241.


Porter LM, Gilbert CC, Fleagle JG.  (2014).  Diet and phylogeny in primate communities.  International Journal of Primatology 35: 1144-1163.


Gilbert CC, Bibi F, Hill A, Beech M.  (2014).  An early guenon from the late Miocene Baynunah Formation, Abu Dhabi, with implications for cercopithecoid biogeography and evolution.  Proceedings of the National Academy of Sciences, USA 111: 10119-10124.


Gilbert CC, Patel BA, Friedman AC, Pugh KD, Fleagle JG, Patnaik R.  (2014).  New Lower Siwalik localities near Ramnagar, India: implications for the earliest Asian great apes and other mammalian lineages.  Special Publication of the Palaeontological Society of India 5: 353-365.  


Rothman JM, Raubenheimer D, Bryer MAH, Takahashi M, Gilbert CC. (2014).  Nutritional contributions of insects to primate diets: implications for primate evolution.  Journal of Human Evolution 71: 59-69.


Roberts P, Delson E, Miracle P, Ditchfield P, Roberts RG, Jacobs Z, Blinkhorn J, Ciochon RL, Fleagle JG, Frost SR, Gilbert CC, Gunnell GF, Harrison T, Korisettar R, Petraglia MD.  (2014).  Continuity of Mammalian Fauna Over the Last 200,000 Years in the Indian Subcontinent.  Proceedings of the National Academy of Sciences, USA 111: 5848-5853.


Rossie JB, Gilbert CC, and Hill A.  (2013).  Early cercopithecid monkeys from the Tugen Hills, Kenya.  Proceedings of the National Academy of Sciences, USA 110: 5818-5822.


Gilbert CC.  (2013).  Cladistic analysis of extant and fossil African papionins using craniodental data.  Journal of Human Evolution 64: 399-433.


Hart JA, Detwiler KM, Gilbert CC, Burrell AS, Fuller JL, Emetshu M, Hart TB, Vosper A, Sargis EJ, Tosi AJ. (2012).  Lesula: A New Species of Cercopithecus Monkey Endemic to the Democratic Republic of Congo and Implications for Conservation of Congo’s Central Basin. PLoS ONE 7(9): e44271. doi:10.1371/journal.pone.0044271


Maiolino S, Boyer DM, Bloch JI, Gilbert CC, and Groenke J.  (2012).  Evidence for a Grooming Claw in a North American Adapiform Primate: Implications for Anthropoid Origins.  PLoS ONE 7(1): e29135. doi:10.1371/journal.pone.0029135


Gilbert CC, Goble ED, Kingston JD, and Hill A.  (2011).  Partial Skeleton of Theropithecus brumpti (Primates: Cercopithecidae) from the Chemeron Formation of the Tugen Hills, Kenya.  Journal of Human Evolution 61: 347-362.


Gilbert CC, Stanley WT, Olson LE, Davenport TRB, and Sargis EJ.  (2011).  Morphological systematics of the kipunji (Rungwecebus kipunji) and the ontogenetic development of phylogenetically informative characters in the Papionini.  Journal of Human Evolution 60: 731-745.


Gilbert CC.  (2011).  Phylogenetic analysis of the African papionin basicranium using 3-D geometric morphometrics: the need for improved methods to account for allometric effects.  American Journal of Physical Anthropology 144: 60-71.


Gilbert CC, Goble ED, and Hill A.  (2010).  Miocene Cercopithecoidea from the Tugen Hills, Kenya.  Journal of Human Evolution 59: 465-483.


Fleagle JG, Gilbert CC, and Baden AL.  (2010).  Primate cranial diversity.  American Journal of Physical Anthropology 142: 565-578.


Gilbert CC and Grine FE.  (2010).  Morphometric variation in the papionin muzzle and the biochronology of the South African Plio-Pleistocene karst cave deposits.  American Journal of Physical Anthropology 141: 418-429.


Gilbert CC, Frost SR, and Strait DS.  (2009).  Allometry, sexual dimorphism, and phylogeny: a cladistic analysis of extant African papionins using craniodental data.  Journal of Human Evolution 57: 298-320.


Gilbert CC, McGraw WS, and Delson E.  (2009).  Brief Communication: Plio-Pleistocene eagle predation on fossil cercopithecids from the Humpata Plateau, southern Angola.  American Journal of Physical Anthropology 139: 421-429.  


Gilbert CC.  (2007).  Identification and description of the first Theropithecus (Primates: Cercopithecidae) material from Bolt's Farm, South Africa.  Annals of the Transvaal Museum 44: 1-10.  


Olejniczak AJ, Gilbert CC, Martin LB, Smith TM, Ulhaas L, and Grine, F.  (2007).  Maxillary molar enamel-dentine junction morphology in anthropoid primates.  Journal of Human Evolution 53: 292-301.  

Gilbert CC and Rossie JB.  (2007).  Congruence of molecules and morphology using a narrow allometric approach.  Proceedings of the National Academy of Sciences, USA 104: 11910-11914.


Gilbert CC.  (2007).  Craniomandibular morphology supporting the diphyletic origin of mangabeys and a new genus of the Cercocebus/Mandrillus clade, Procercocebus.  Journal of Human Evolution 53: 69-102.  


Patel BA, Gilbert CC, and Ericson KE.  (2007).  Cercopithecoid cervical vertebral morphology and implications for the presence of Theropithecus in early Pleistocene Europe.  Journal of Human Evolution 52: 113-129.  


Gilbert CC.  (2005).  Dietary ecospace and the diversity of euprimates during the Early and Middle Eocene.  American Journal of Physical Anthropology 126: 237-249.


Pochron ST, Fitzgerald J, Gilbert CC, Lawrence D, Grgas M, Rakotonirina G, Ratsimbazafy R, Rakotosoa R, and Wright PC.  (2003).  Patterns of female dominance in Propithecus diadema edwardsi of Ranomafana National Park, Madagascar.  American Journal of Primatology 61: 173-185.

Edited Book Chapters


Frost SF, Sena C, and Gilbert CC (accepted, in press).  Plio-Pleistocene cercopithecoid evolution and biogeography: a contemporaneous comparison with Paranthropus. In: Constantino PJ, Reed KE, Wood BA (Eds.), The Forgotten Lineage(s): Paelobiology of Paranthropus


Gilbert CC and Hill A.  (in press).  Chapter 13: Primates of the late Miocene Baynunah Formation, Abu Dhabi.  In: Bibi F, Hill A, Beech MJ, Kraatz B (Eds.), Sands of Time: Ancient Life in the Late Miocene of Abu Dhabi, United Arab Emirates. Dordrecht: Springer.  pp. 203-218. 


Frost SF, Gilbert CC, and Nakatsukasa M (2022).  The colobine fossil record.  In: Matsuda I, Grueter CC, Teichroeb JA (Eds.), The Colobines: Natural History, Behaviour and Ecological Diversity. Cambridge: Cambridge University Press.  pp. 13-31.


Frost SF, Gilbert CC.  (2021).  Entry for “Old World Monkeys”.  eLS (Encyclopedia of Life Sciences).  DOI: 10.1002/047001590X


Gilbert CC, Pugh KD, and Fleagle JG.  (2020).  Chapter 17: Dispersal of Miocene hominoids (and pliopithecoids) from Africa to Eurasia in light of changing tectonics and climate.  In: Prasad GVR, Patnaik R (Eds.), Biological Consequences of Plate Tectonics: New Perspectives on Post-Gondwanal and Break-up- A Tribute to Ashok Sahni. Cham: Springer.  pp. 393-412.

Baden AL and Gilbert CC.  (2016).  Entry for Propithecus coquereli.  In: All the World’s Primates, Noel Rowe, Marc Myers, eds.  Primate Conservation Inc., Charlestown RI.


Fleagle JG, Royer DF, and Gilbert CC.  (2016).  Entry for Homo sapiens.  In: All the World’s Primates, Noel Rowe, Marc Myers, eds.  Primate Conservation Inc., Charlestown RI.

Fleagle JG and Gilbert CC.  (2011).  Introduction to All the World’s Primates: Primate Evolution.  In: All the World’s Primates, Noel Rowe, Marc Myers, eds.  Primate Conservation Inc., Charlestown RI.


Jernvall J, Gilbert CC, and Wright, PC.  (2008).  Peculiar teeth homologies of the greater bamboo lemur (Prolemur = Hapalemur simus): when is a paracone not a paracone?  In: Elwyn Simons: A Search for Origins, Fleagle JG and Gilbert CC, eds.  New York: Springer.  pp. 335-342.


Simons EL, Chatrath P, Gilbert CC, and Fleagle JG.  (2008).  Five Decades in the Fayum.  In: Elwyn Simons: A Search for Origins, Fleagle JG and Gilbert CC, eds.  New York: Springer.  pp. 51-70.


Fleagle JG and Gilbert CC.  (2006).  Biogeography and the primate fossil record: the role of tectonics, climate, and chance.  In: Primate Biogeography, Lehman S and Fleagle JG, eds.  New York: Springer.  pp. 375-418.

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